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Embryonic stem cells require special conditions in order to be maintained in their pluripotent state. Early culture methods relied upon plating ES cells upon a feeder layer, usually of mouse embryonic fibroblasts. Later, it was determined that these feeder cells provided a factor, LIF, that blocked differentiation of ES cells. This has led to the development of feeder-free ES culture methods that can sometimes be used instead of a feeder cell layer.
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